The coconut palm, Cocos nucifera
, is an erect palm in the family Arecaceae which is grown its fruits, used primarily for the extraction of coconut oil for use in cooking. The coconut palm has an erect or slightly curved stem which grows from a swollen base. The stem is smooth, light gray in color and has prominent leaf scars. The stem is topped with a crown of 60–70 spirally arranged leaves. The leaves are long (up to 7 m/23 ft), pinnately divided and composed of 200–250 tapering leaflets. The inflorescence is a spike produced at the leaf axil with 20–60 branches, each with a female flower at the base and many male flowers. The fruit is a drupe containing a single seed. It is ovoid in shape with three sides divided by ridges. The exocarp and the mesocarp make up the husk of the coconut. The seed is protected by a thick, stony shell, or endocarp, and is partially filled with a liquid known as coconut water. The edible endosperm is white and meaty and can be between 1.0 and 2.5 cm (0.4–1.0 in) thick. Coconut palms can reach a height of 30 m (98 ft), produce up to 75 fruits a year, and live for up to 90 years. The origin of the coconut is unknown although the center of genetic diversity lies in Southeast Asia.
Harvested coconut drupes
Coconut seed cut open to reveal flesh
Coconut palms can reach 30 m (98 ft)
Close-up image of coconut palm leaf
Coconut palms with coconuts
Coconuts are primarily cultivated for their oil. The oil is extracted from the endosperm and is used in cooking. Low grade oils are used in the production of soaps. The endosperm can be consumed fresh or dry and is commonly grated for use in baking. Young coconuts, termed waternuts, are commonly sold in tropical resorts, where the liquid is drunk directly from the shell. Coconuts may also be used to produce coconut milk by squeezing the liquid from the grated endosperm. Coconut milk is popularly used in many dishes from Southeast Asia.
The coconut palm is a tropical plant and is generally grown in humid, tropical regions. It grows optimally in areas with an annual mean temperature of 27°C (80.6°F) with in excess of 2000 hours of sunlight per year. The palm will thrive in a wide range of soils from sand to clay as long as they are well draining and well aerated with a pH between 4.3 and 8.0. Although palms are often found growing on sandy beaches, they can be successfully grown inland but will not tolerate freezing temperatures.
Coconut palms are propagated exclusively from seed. The seeds are ready for planting when the coconut milk can be heard sloshing around inside the seed when it is shaken. Seeds are germinated by planting in seed beds before transferring to a polythene bag or nursery beds after germination. Seeds should be planted on their sides in a shallow hole with enough soil to cover about one third of the seed. The seed should be watered regularly to prevent it drying out. Germination usually occurs after about 3 months but may take up to 6 months.
Coconut seedling can be transplanted from 6 months onwards or transferred to pots and grown further in the nursery. Trees require a wide spacing and are typically planted 8–9 m (26–30 ft) apart allowing a further 8–9 m (26–30 ft) between rows. Dwarf varieties can be spaced closer together and are typically planted 7.5 m (25 ft) apart allowing another 7.5 m (25 ft) between rows
Broschat, T. K. & Crane, J. H. (2011). The Coconut Palm in Florida. University of Florida IFAS Extension. Available at: http://edis.ifas.ufl.edu/pdffiles/MG/MG04300.pdf. [Accessed 13 November 14]. Free to access
CABI Crop Protection Compendium. (2014). Cocos nucifera datasheet. Available at: http://www.cabi.org/cpc/datasheet/11788. [Accessed 13 November 14]. Paid subscription required
Chan, E. & Elevitch, C. R. (2006). Cocos nucifera
(coconut). Species profiles for Pacific Agroforestry. Traditional Tree Initiative. Available at: http://www.agroforestry.net/images/pdfs/Cocos-coconut.pdf. [Accessed 13 November 14]. Paid subscription required
Duke, J. A. (1983). Cocos nucifera
L.. Handbook of Energy Crops. Available at http://www.hort.purdue.edu/newcrop/duke_energy/Cocos_nucifera.html#Cultivation. [Accessed 13 November 14]. Free to access
Elliott, M. L., Broschat, T. K., Uchida, J. Y. & Simone, G. W. (2004). Compendium of Ornamental Palm Disease and Disorders. American Phytopathological Society Press. Available at: http://www.apsnet.org/apsstore/shopapspress/Pages/43143.aspx. Available for purchase from APS Press
Common Pests and Diseases
Category : Fungal
Bud rot and nutfall
Coconut tree infected with bud rot pathogen Phytophthora palmivora
Chlorosis of youngest open leaves; leaves rapidly turning necrotic; necrotic spots on leaf bases; unopened spear leaves can be pulled away from the plant easily; removal of unopened spear leaves reveals soft, pink-red tissue with foul smell; leaf necrosis spreading through central crown leaves; woody parts of plant may have water-soaked, pink lesions with dark borders; infected inflorescences abort nuts.
Control of the disease is reliant on good sanitation practices and the use of appropriate systemic fungicides; remove all infected debris and dead trees from plantation and destroy; irrigate trees early in the day to allow surfaces to dry off during the day.
Ganoderma butt rot
Close-up of Ganoderma root and butt rot fruiting bodies
Ganoderma root and butt rot (Ganoderma australe) fruiting bodies
Older fronds turning yellow and gradually wilting and drooping; fronds collapsing and dying; internal tissue of lower stem discolored; overall reduction in vigor.
Spacing trees widely limits the chance of infection through root grafts; avoid damaging tree trunks with tools and machinery; remove any dead or severely damaged trees from plantation immediately, including any that have been killed by natural processes; if a site is known to be infected with the disease, the ground should be fallowed for at least 1 year prior to a new plantation being established.
Gray leaf blight
Pestalotiopsis leaf spot (Pestalotiopsis palm arum) sign
Small, yellow-brown spots on leaflets which develop gray centers and dark green borders; lesions coalesce to form large necrotic patches; tips of leaflets turning gray; canopy has blighted appearance.
Disease usually only merits control in coconut nurseries as infection of mature coconut palms are rare; appropriate broad spectrum protective fungicides should be applied.
Lethal bole rot
Oldest leaves of palm turning yellow and wilting; reddish-brown rot in bole tissue; destruction of root system
Any infected trees must be uprooted and burned; area can only be replanted once soil is treated for the disease
Stem bleeding disease
Chalara paradoxa (syn. Thielaviopsis paradoxa, Ceratocystis paradoxa)
The canopy of the coconut in the center is wilted and necrotic due to a trunk infection by Thielaviopsis paradoxa.
Trunk collapse due to stem bleeding disease
The three coconuts on the left have died from Thielaviopsis trunk rot. The palm in the foreground exhibits trunk collapse.
The trunk of this coconut was just beginning to exhibit "stem bleeding", but the large rusty-brown area at the top was already soft.
Only one side of this trunk has significant rot due to Thielaviopsis paradoxa. The fungus rots the trunk tissue from the outside to the inside.
Stem bleeding on a coconut trunk. The top of the blackened area was very soft and could be easily pushed in with the fingers.
Soft, yellow rot on trunk; affected areas are dark and turn black as they mature; a reddish-brown liquid may ooze from rotting regions and spill down trunk.
Avoid wounding palms with machinery and tools to reduce disease incidence; disease can be controlled with applications of the fungicide benomyl where registered; infected trees should be removed and destroyed as soon as possible.
Category : Viral
Coconut cadang-cadang viroid (CCCVd)
Area with cadang-cadang disease showing trees in the early, medium and late stages of the disease.
Newly formed nuts more rounded than in previous years; nuts exhibit scarring on the surface; chlorotic spots on leaves; stunted inflorescences with tip necrosis; leaves begin to decline in size and number; death of palm.
There is currently no known method of controlling the disease.
Coconut foliar decay
Coconut foliar decay virus (CFDV)
Some varieties which have the disease may show no symptoms, others exhibit partial yellowing of leaves which begins to spread to leaf tip; necrosis of petioles causing leaves to die and hang from palm canopy
Category : Other
Lethal yellowing (Palm lethal yellowing)
Candidatus Phytoplasma palmae (PLY)
Damage due to lethal yellowing
Cocos nucifera spear leaf is dying just as the last leaves are discoloring due to lethal yellow phtyoplasma
Infected coconut trees
Lethal yellowing infected palm
Symptoms of lethal yellowing on coconut palm
Lethal yellowing damaged tree
Lethal yellowing infected coconut trees
Symptoms of lethal yellowing on coconut palm
Premature dropping of fruit; fruit with brown-black water-soaked appearance; necrosis of inflorescences; flower stalks turn black; lower, older leaves turning yellow; entire crown turning yellow; yellow leaves turn brown, dry out and hang from canopy.
The most effective method of managing the disease is to plant resistant coconut varieties such as Malayan dwarf or Maypan; antibiotic treatment is effective but not usually practical for large scale plantings.
Category : Insects
Damged and/or aborted flowersd; sunken necrotic lesions and scars on nuts; young nuts may exude gum (gummosis) and die; many nuts fall from tree; adult insect is a brown-red with well-developed wings; nymphs are brown-red or green in color with long antennae and feed at the calyx of the nut
Natural enemies of the coconut bug include weaver ants, conserve bushes and trees around plantation which are habitats for weaver ants or intercrop with mango, guava or citrus which are attractive to weaver ants; connect canopy with ropes or sticks to allow weaver ants to move between trees
Omiodes blackburni (earlier Hedylepta blackburni)
Coconut leafroller (Hedylepta blackburni) damage to coconut plant
Damage due to coconut leafrollers
Coconut leafroller pupa, larva and frass.
Coconut leafroller (Hedylepta blackburni) adult
Coconut leafroller larvae
Coconut leafroller pupae and frass
Coconut leafroller feeding on leaves
Coconut leafroller (Hedylepta blackburni) larvae damage to coconut leaves.
Coconut leafroller rolls leaf near midrib of frond
The larvae feeds on under surface of leaves. Usually they found protected by silken web. Initially they feed on the lower epidermis leaving the upper epidermis intact. The larvae often fold two sides of leaflets by a silken thread and feed inside. The later stage larvae feeds on both upper and lower epidermis of leaves. Severe infestation leads to skeletonization of fronds.
Encourage natural enemies in the orchard.
Coconut rhinoceros beetle
Damaged coconut frond
coconut rhinoceros beetle (Oryctes rhinoceros) adult
Lure and trap for Coconut Rhinoceros Beetle
coconut rhinoceros beetle injury to young coconut tree
coconut rhinoceros beetle (Oryctes rhinoceros) pupae
Damage due to coconut rhinoceros beetle
Coconut rhinoceros beetle larvae
Coconut Rhinoceros Beetle (CRB)
coconut rhinoceros beetle (Oryctes rhinoceros) injured trees
Damage due to rhinoceros beetle
Rhinoceros beetle damage
The Asiatic rhinoceros beetle or coconut rhinoceros beetle (Oryctes rhinoceros) damage to coconut
V-shaped cuts in palm fronds or holes in leaf midribs caused by beetles boring into crown to feed; adult insect is a large black beetle with a curved spine on its head; larvae are creamy white grubs with brown heads and 3 sets of prolegs at the anterior (head) end.
Destroy any decaying logs in plantation by chopping and burning to kill any larvae that may be inside; remove any dead trees from plantation and destroy by burning; plant a cover crop to deter egg laying by females as they do not lay eggs in areas covered by vegetation; hooked wire can be used to extract larvae that are boring into young crowns.
coconut scale (Aspidiotus destructor) infestation
coconut scale (Aspidiotus destructor)
Scale on coconut foliage
Coconut scale (Aspidiotus destructor) adults
coconut scale (Pinnaspis buxi)
Coconut scale damage
coconut scale (Aspidiotus destructor) adults and early instars
Coconut scale infestation
Coconut scale damage
Pale yellow spots on leaves; entire leaves yellowing; leaves turning brown and dropping prematurely; adult insect is a flattened oval, resembling a scale, which is red-brown in color.
May be possible to control coconut scale by pruning infested parts of trees and destroying by burning; chemical control may be necessary.
Mealybugs (Pineapple mealybug, Striped mealybug, Cocoa mealybug, etc.)
Coconut mealybugs (Nipaecoccus nipae)
Coconut (Cocos nucifera): Mealybugs and scales on leaflet
Coconut mealybug damage
Colony of coconut mealybugs (Nipaecoccus nipae)
Adults of coconut mealybug (Nipaecoccus nipae)
coconut mealybugs (Nipaecoccus nipae) tended by ants
Coconut mealybug (Nipaecoccus nipae) adult
Flattened oval to round disc-like insect covered in waxy substance on tree branches; insects attract ants which may also be present; insect colony may also be associated with growth of sooty mold due to fungal colonization of sugary honeydew excreted by the insect; symptoms of direct insect damage not well documented but trees may exhibit symptoms of cocoa swollen shoot (see disease entry).
Mealybugs can potentially be controlled by natural enemies such as lady beetles but are commonly controlled using chemicals; chemical pesticides may also decrease populations of natural enemies leading to mealybug outbreaks.
Category : Mites
Eriophyid coconut mite
Mite feeding injury
Coconut mite damage coconut fruits
Symptoms due to coconut mite infestation
Coconut: Eriophyid mites injury to nuts
Scarring of coconut fruits due to coconut mites
Eriophyid mites feeding injury to coconut
Coconut mite feeding injury
Coconut mite damage on fruits of coconut
The mites suck sap from young nuts. Generally they feed on meristematic zone, i.e., the area which is covered by perianth. The infestation starts very early. As the nut develops the feeding leaves brown fissures that extending down from the perianth. The nut becomes small and distorted.
Provide proper fertilizer and water for trees to withstand mite damage. Encourage natural enemies of mite in the orchard. If infestation is severe, apply suitable insecticide by root feeding or stem injection.
Category : Nematodes
Red ring nematode (Coconut palm nematode)
Nuts falling prematurely; withering inflorescences; yellowing leaves which then turn brown; orange to red-brown ring of discoloration when a cross section is taken of lower stem.
If a tree becomes infected it should be removed and destroyed; control of the disease is currently limited to efforts to control the weevil which transmits the nematode to the palms.